ANALISIS AKTIVITAS STOMATA PADA TUMBUHAN COCOR BEBEK (Kalanchoe pinnata (Lam) Pers. ) DENGAN ADAPTASI CAM (Crassulacean Acid Metabolism)
DOI:
https://doi.org/10.30998/p0nwwv73Keywords:
Stomata, Kalanchoe pinnata, CAM, FotosintesisAbstract
Kalanchoe pinnata (Lam) Pers., commonly known as Cocor Bebek, is a succulent plant capable of surviving in arid environments through a physiological adaptation known as Crassulacean Acid Metabolism (CAM). The aims of this research is to describe the stomatal activity of the cocor bebek plant which one of the species with a CAM (Crassulacean Acid Metabolism) photosynthesis pathway. This research used descriptive qualitative method with purposive sampling. The method is observing stomatal activity were conducted on the lower leaf surface using a clear nail-polish replica technique and a light microscope every three hours. The results show that the stomata of Kalanchoe pinnata remain mostly closed during the daytime and open at night. The widest stomatal opening occurred at 21:00, with a higher number of open stomata compared to other observation times. This activity is one of the physiological adaptation of CAM plants, which close their stomata during the morning until afternoon day to reduce water loss and open them at night for carbon fixation. The stomata identified were anisocytic and predominantly located on the abaxial leaf surface. This study confirms that the opening and closing pattern of the stomatal of cocor bebek is closely related to water use efficiency mechanism characteristic in CAM plants.
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Alhafidz, N.N.K, Zahirah, N.L, Azzahra, T.A, Wulaningsih, R.D. (2025). Analisis Karakteristik Stomata Daun Rhoeo discolor Berdasarkan Waktu Pengambilan Yang Berbeda. Jurnal Biologi Science and Education, 14 (1), 45-53
Asriyani, L. (2017). Identifikasi Penentuan Waktu Optimal Pembukaan Stomata Alang-Alang (Imperata cylindrica L.) di UIN Raden Intan (Studi Deskriptif Sebagai Sumber Belajar Peserta Didik Materi Fotosintesis SMA Kelas XII Semester Ganjil) . Skripsi. Lampung : Universitas Islam Negeri Raden Intan.
Aulia, O. M., Amintarti, S., & Rezeki, A. (2023). Tipe-Tipe Stomata Tumbuhan Myrtaceae Di Lingkungan Kampus Fkip Ulm Sebagai Booklet Bahan Ajar Pendamping Mata Kuliah Anatomi Tumbuhan. BIOEDUKASI (Jurnal Pendidikan Biologi), 14(2), 230. https://doi.org/10.24127/bioedukasi.v14i2.7791
Bharath, P., Gahir, S., & Raghavendra, A. S. (2021). Abscisic Acid-Induced Stomatal Closure: An Important Component of Plant Defense Against Abiotic and Biotic Stress. Frontiers in Plant Science, 12(March), 1–18.
https://doi.org/10.3389/fpls.2021.615114
Campbell, N.A., Reece, J.B., & Mitchell, L.G. (2002). Biologi. Jilid 1. Edisi Kelima. Alih Bahasa: Wasmen. Jakarta: Penerbit Erlangga.
Fadli, M.R. (2021). Memahami desain Metode Penelitian Kualitatif. Jurnal Humanika Kajian Ilmiah Mata Kuliah Umum, 21(1), 35-54 https://doi.org/10.21831/HUM.V21I1.38075
Fauziah, A., & ‘Izzah, A. S. Z. (2019). Analisis Tipe Stomata Pada Daun Tumbuhan. Prosiding Seminar Nasional HAYATI VII Tahun 2019, 35–39.
Gilman, I.S & Erika J. S. (2020) Crassulacean Acid Metabolism. Curent Biology Magazine https://www.cell.com/current-biology/fulltext/S0960-9822(19)31575-1
Gotoh, E., Oiwamoto, K., Inoue, S. I., Shimazaki, K. I., & Doi, M. (2019). Stomatal response to blue light in crassulacean acid metabolism plants Kalanchoe pinnata and Kalanchoe daigremontiana. Journal of Experimental Botany, 70(4), 1367–1374. https://doi.org/10.1093/jxb/ery450
Hidayat, Estiti. 1995. Anatomi Tumbuhan Berbiji. Bandung : ITB
Lim, et al. (2025) Synthetic Crassulacean Acid Metabolism (SynCAM) For Improving Water Use Efficiency In Plants. Journal Of Biological Sciences Vol 380.
Muthi’ah, S. N. (2022). Identifikasi dan Karakterisasi Tipe Stomata Pada Hibiscus rosa-sinensis, Tamarindus indica, dan Mangifera indica Dengan Teknik Replika . Indigenous Biologi : Jurnal Pendidikan Dan Sains Biologi, 5(1),9–14. https://doi.org/10.33323/indigenous.v5i1.295
Oktaviani, E., & Daningsih, E. (2022). Distribusi dan Luas Stomata pada Tanaman Hias Monokotil. Jurnal Ilmu Pertanian Indonesia, 27(1), 34–39. https://doi.org/10.18343/jipi.27.1.34
Purwanti R. & Diah, N.K. (2025). Karakteristik Simplisia Dan Skrining Fitokimia Senyawa Alkoloid, Flavonoid, Tanin, Dan Saponin Ekstrak Etanol Daun Cocor Bebek (Kalanchoe pinnata (Lam.) Pers). Jurnal Permata Indonesia, 17 (1), 1-8. https://doi.org/10.59737/jpi.v17i1.346
Prastika, D., Sarjani, T. M., Mahyuni, S. R., Hariani, I., Ramadhan, D. A., Rezeki, S., Tiara, R., Hendrik, E., Aulia, R., & Amalia, T. (2023). Identifikasi Tipe Stomata Anggota Suku Myrtaceae di Kota Langsa. Jurnal Sains Dan Edukasi Sains, 6(1), 20–27. https://doi.org/10.24246/juses.v6i1p20-27
Salira, T., & Chatri, M. (2021). Tipe Stomata Pada Beberapa Tanaman Dari Familia Rubiaceae. Prosiding Semnas Bio, 2013, 1–5.
Santos, M. G., Davey, P. A., Hofmann, T. A., Borland, A., Hartwell, J., & Lawson, T. (2021). Stomatal Responses to Light, CO2, and Mesophyll Tissue in Vicia faba and Kalanchoë fedtschenkoi. Frontiers in Plant Science, 12(October), 1–17. https://doi.org/10.3389/fpls.2021.740534
Saputri dan Eka. 2017. Pola pembukaan dan Penutupan Stomata Pada Tiga Spesies Anggota Genus Sansivieria. Prosiding Seminar Nasional Pendidikan. ISBN : 978-602-70313-2-6 Universitas Muhamadiyah Metro https://repository.ummetro.ac.id/files/artikel/634471684febe3f0ae0152fd49daa5c3.pdf
Schiller K. & Andrea B (2021) Engineering Of Crassulacean Acid Metabolism. Annual Review Of Plant Biology, Vol 72 Hal 77-103 , https://doi.org/10.1146/annurev-arplant-071720-104814
Singh, R., Ajay, G., Khushboo, S. (2022). Botanical description, Photochemistry, Traditional Uses, and Pharmacology of the “Wonder Plant” Kalanchoe Pinnata (Linn.) Journal of Pharmaceutical and Medicinal Chemistry, 8(1), 11-24 . http://dx.doi.org/10.21088/jpmc.2395.6615.8122.1
Suyatman, S. (2021). Menyelidiki Energi Pada Fotosintesis Tumbuhan. INKUIRI: Jurnal Pendidikan IPA, 9(2), 134. https://doi.org/10.20961/inkuiri.v9i2.50085
Widiastuti, Ni Komang. (2023). Tanaman Obat Tradisional ; Cocor Bebek (Kalanchoe pinnata). https://www.scribd.com/document/622524594/BUKU-COCOR-BEBEK
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